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21 Aug, 2019
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Argiles Et Grès à Reptiles Formation

Argiles et Grès à Reptiles Formation (meaning Reptile Clay and Sandstone) also known as the Argiles Rutilantes Formation is an early Maastrichtian French geologic formation in the département of Var preserving the remains of several types of dinosaurs and other extinct organisms.

Fossil content

An abelisaurid similar to Arcovenator is known from Département des Bouches-du-Rhone, while enantiornithean and indeterminate avialan remains are known from Département du Var.

Dinosaurs

Dinosaurs of the Grès à Reptiles
Genus	Species	Location	Stratigraphic position	Abundance	Notes	Images
Abelisauridae	Indeterminate	Bouches du Rhône	Lower		A large abelisaurid that was similar to Arcovenator.
Arcovenator	A. escotae	Var	Lower		A large abelisaurid.
Atsinganosaurus	G. meridionalis	Bouches du Rhône	Lower		A titanosaur Lirainosaurinae.
Castignovolucris	C. sebei	Castigno valley, Villespassans, Hérault	Lower (i.e. continental red clays)		An enantiornithine bird.
Garrigatitan	A. velauciensis	Bouches du Rhône	Lower		A titanosaur Lirainosaurinae.
Gargantuavis	G. philoinos	Var
Hypselosaurus	H. priscus	Bouches-du-Rhône Var			Actually indeterminate sauropod remains.
Matheronodon	M. provincialis	Bouches-du-Rhône			A rhabdodontid ornithopod
Theropoda	Indeterminate	Var			Previously referred to "Megalosaurus" pannoniensis
Pyroraptor	P. olympius	Bouches-du-Rhône		"Fragmentary postcrania and teeth"
Rhabdodon	R. priscus	Bouches-du-Rhône Var
Rhodanosaurus	R. lugdunensis	Bouches-du-Rhône
Variraptor	V. mechinorum	Var Bouches-du-Rhône

Mammals of the Grès à Reptiles
Genus	Species	Location	Stratigraphic position	Abundance	Notes	Images
Mistralestes	M. arcensis	La Cairanne
Valentinella	V. vitrollense	Vitrolles-La Plaine

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Crocodylians

Turtles

Pterosaurs

Mistralazhdarcho maggii

Fish

Fossil eggs

Flora

References

^ Tabuce et al., 2013
^ Weishampel et al., 2004, pp.588-593
^ Argiles et Grès à Reptiles Formation at Fossilworks.org
^ Tortosa, Thierry; Buffetaut, Eric; Vialle, Nicolas; Dutour, Yves; Turini, Eric; Cheylan, Gilles (2014-01-01). "A new abelisaurid dinosaur from the Late Cretaceous of southern France: Palaeobiogeographical implications". Annales de Paléontologie. 100 (1): 63–86. Bibcode:2014AnPal.100...63T. doi:10.1016/j.annpal.2013.10.003. ISSN 0753-3969.
^ Díez Díaz, Verónica; Garcia, Géraldine; Pereda Suberbiola, Xabier; Jentgen-Ceschino, Benjamin; Stein, Koen; Godefroit, Pascal; Valentin, Xavier (2020-12-21). "A new titanosaur (Dinosauria: Sauropoda) from the Upper Cretaceous of Velaux-La-Bastide Neuve (southern France)". Historical Biology. 33 (11): 2998–3017. doi:10.1080/08912963.2020.1841184. ISSN 0891-2963. S2CID 234404741.
^ Buffetaut, E.; Angst, D.; Tong, H. (2023). "A new enantiornithine bird from Upper Cretaceous non-marine deposits at Villespassans (Hérault, southern France)". Annales de Paléontologie. 109 (1). 102585. Bibcode:2023AnPal.10902585B. doi:10.1016/j.annpal.2022.102585. S2CID 257863645.
^ B. Vila; A. Galobart; J.U. Canudo; J. Le Loeff (2012). "The diversity of sauropod dinosaurs and their first taxonomic succession from the latest Cretaceous of southwestern Europe: Clues to demise and extinction". Palaeogeography, Palaeoclimatology, Palaeoecology. 350–352 (15): 19–38. Bibcode:2012PPP...350...19V. doi:10.1016/j.palaeo.2012.06.008.
^ "14.8 Département du Var, France; 2. Grès à Reptiles Formation," in Weishampel et al., 2004, p.589
^ "14.8 Département du Var, France; 2. Grès à Reptiles Formation" and "14.38 Département des Bouches-du-Rhone, France; 1. Grès à Reptiles Formation," in Weishampel et al., 2004, pp.589-590
^ "14.38 Département des Bouches-du-Rhone, France; 1. Grès à Reptiles Formation," in Weishampel et al., 2004, p.590
^ "Table 10.1," in Weishampel et al., 2004, p.199
^ Martin, 2007
^ Blanco & Brochu, 2016
^ Le Loeuff & Buffetaut, 1998
^ Tong et al., 1998
^ Martin et al., 2016
^ Vullo et al., 2018
^ Viany-Liaud et al., 1994

Bibliography

Vullo, R.; Garcia, G.; Godefroit, P.; Cincotta, A.; Valentin, X. (2018), "Mistralazhdarcho maggii, gen. et sp. nov., A New Azhdarchid Pterosaur from the Upper Cretaceous of Southeastern France", Journal of Vertebrate Paleontology, 38 (4): 1–16, doi:10.1080/02724634.2018.1502670
Blanco, A.; Brochu, C. A. (2016), "Intra- and interspecific variability in allodaposuchid crocodylomorphs and the status of western European taxa", Historical Biology, 29: 395–408
Martin, J. E.; Delfino, M.; Garcia, G.; Godefroit, P.; Berton, S.; Valentin, X. (2016), "New specimens of Allodaposuchus precedens from France: intraspecific variability and the diversity of European Late Cretaceous eusuchians", Zoological Journal of the Linnean Society, 176 (3): 607–631, doi:10.1111/zoj.12331
Tabuce, Rodolphe; Tortosa, Thierry; Vianey-Liaud, Monique; Garcia, Géraldine; Lebrun, Renaud; Godefroit, Pascal; Dutour, Yves; Berton, Sévérine; Valentin, Xavier; Cheylan, Gilles (2013), "New eutherian mammals from the Late Cretaceous of Aix-en-Provence Basin, south-eastern France", Zoological Journal of the Linnean Society, 169 (3): 653–672, doi:10.1111/zoj.12074, ISSN 0024-4082
Martin, J. E (2007), "New material of the Late Cretaceous globidontan Acynodon iberoccitanus (Crocodylia) from southern France", Journal of Vertebrate Paleontology, 27 (2): 362–372, doi:10.1671/0272-4634(2007)27[362:NMOTLC]2.0.CO;2
Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (2004), The Dinosauria, 2nd edition, Berkeley: University of California Press, pp. 1–880, ISBN 0-520-24209-2, retrieved 2019-02-21
Le Loeuff, J.; Buffetaut, É. (1998), "A new dromaeosaurid theropod from the Upper Cretaceous of southern France.", Oryctos, 1: 105–112
Tong, H.; Gaffney, E. S.; Buffetaut, É. (1998), "Foxemys, a new side-necked turtle (Bothremydidae: Pelomedusoides) from the Late Cretaceous of France", American Museum Novitates (3251): 1–19
Vianey-Liaud, M.; Mallan, P.; Buscail, O.; Montgelard, C. (1994), Review of French dinosaur eggshells: morphology, structure, mineral, and organic composition. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge, pp. 151–183