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21 Aug, 2019
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Cerro Del Pueblo Formation

The Cerro del Pueblo Formation is a geological formation in Coahuila, Mexico, whose strata date back to the latest Campanian of the Late Cretaceous, just before the Campanian-Maastrichtian boundary. Dinosaur remains are among the fossils that have been recovered from the formation. The formation is believed to correlate with the Baculites reesidesi and Baculites jenseni ammonite zones, which dates it to 73.63-72.74 Ma.

The Cerro del Pueblo Formation includes marine, brackish, and non-marine environments, and was laid down in a coastal lowland environment, with Coahuila being on the edge of the Western Interior Seaway at the time. Fossils are abundant in the Cerro del Pueblo Formation, and include dinosaurs and other vertebrates, plants and seeds, and invertebrates including oysters, snails, and other molluscs. The formation is underlain by the Parras Shale and is overlain by the Cerro Huerta Formation.

Dinosaurs

Ornithischians

Remains of the following ornithischians have been found in the formation:

Ankylosaurs

Ankylosaurs of the Cerro del Pueblo Formation
Genus	Species	Material	Notes	Images
Ankylosauria	Indeterminate	A femur fragment, phalanx and a possible osteoderm.	Indeterminate ankylosaur remains.
Ankylosauridae	Indeterminate	Osteoderms.	Indeterminate ankylosaurid remains.
Nodosauridae	Indeterminate	A caudal vertebra, osteoderms and a tooth.	Indeterminate nodosaurid remains.

Ceratopsians

Ceratopsians of the Cerro del Pueblo Formation
Genus	Species	Material	Notes	Images
Centrosaurinae	Indeterminate	A partial right squamosal.	Probably represents a new taxon.
Ceratopsidae	Indeterminate	Squamosal fragment, metatarsal, metacarpal, sacral vertebrae, proximal end of femora, proximal end of ulna, possible frill fragments and teeth.	Indeterminate ceratopsid remains.
Chasmosaurinae	Indeterminate	Supraorbital horncores, an orbit, dorsal vertebra, indeterminate cranial fragments, limb bones, and unspecified postcranial elements pertaining to juvenile specimens.	Indeterminate chasmosaurine remains belonging to juvenile specimens.

Ornithopods

Ornithopods of the Cerro del Pueblo Formation
Genus	Species	Material	Notes	Images
Coahuilasaurus	C. lipani		A kritosaurin hadrosaurid
Hadrosauridae	Indeterminate	A braincase, postorbital, quadrate, laterosphenoid, dentaries, mandibles, maxillae, cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, neural spines, ribs, ilium, femora, fibulae, tibiae, humeri, ulnae, metatarsals, metacarpals, pedal phalanxes, pedal phalanges, pedal ungula, phalanges, phalanxes, various bone fragments, fibulae, scapulae, radii, coracoids, astragali, indeterminate pelvic bones, ossified tendons, isolated teeth and integumentary impressions.	Indeterminate hadrosaurid remains from numerous specimens.
Kritosaurus	K. navajovius	Predentary, symphyseal processes, rostral portion of dental battery of dentaries, rostral region of premaxillae, and fragments of maxilla.	A kritosaurin hadrosaurid, originally referred to K. navajovius, but reclassified to Coahuilasaurus.
Lambeosaurinae	Indeterminate	Maxillae, premaxillae, quadrate, partial braincase, dentaries, cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, neural spines, ribs, ilia, ischium, femora, fibulae, tibiae, humeri, ulnae, metatarsals, metacarpals, pedal phalanges, phalanges, pedal unguals, astragali, radii, scapulae, pubes and other unprepared elements.	Indeterminate lambeosaurine remains from numerous specimens, including a large hadrosaurid.
Latirhinus	L. uitstlani	Caudal vertebrae, scapulae, humeri, ulnae, coracoid, metacarpals, manual ungual, ilia, ischia, femora, tibiae, fibula, and astragalus.	Holotype specimen is chimeric as it consists of multiple individuals. Some material from the holotype specimen is assigned to Lambeosaurinae indet. and Saurolophinae indet..
Saurolophinae	Indeterminate	A braincase, brain endocast, dentaries, maxillae, cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, chevrons, neural spines, ribs, ilium fragments, femora, fibulae, tibiae, humeri, ulnae, metatarsals, phalanges, phalanxes, scapulae, a possible coracoid, radii, ossified tendons, and undescribed juvenile specimens.	Indeterminate saurolophine remains from numerous specimens, including juvenile individuals.
Tlatolophus	T. galorum	An almost complete skull, mandible, partial ilium, partial ischium, a femur, scapula, coracoid, sacral vertebrae, caudal vertebrae, and chevrons.	A parasaurolophin hadrosaur, closely related to Charonosaurus and Blasisaurus.
Velafrons	V. coahuilensis	Dentaries, predentaries, ceratobranchial, premaxillae, maxillae, jugal, quadrate, nasal, skull roof, cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, neural arches, cervical ribs, ribs, chevrons, scapula, coracoids, humeri, ulnae, radii, metacarpals, manual phalanges, ilium, ischia, pubes, femora, tibiae, astragalus, metatarsals and pedal phalanges.	A lambeosaurine hadrosaurid known only from a juvenile specimen.

Thescelosaurids

Thescelosaurids of the Cerro del Pueblo Formation
Genus	Species	Material	Notes	Images
Thescelosauridae	Indeterminate	A right premaxillary tooth and a posterior sacral vertebral centrum.	Indeterminate thescelosaurid remains.

Saurischians

Remains of the following saurischians have been found in the formation:

Theropods

Theropods of the Cerro del Pueblo Formation
Genus	Species	Material	Notes	Images
Caenagnathidae		3 tibia fragments, distal, proximal and medial shaft elements respectively.	An oviraptorosaur. Two different morphotypes are suggested, a small one, less than a meter in length. And a big morphotype, almost reaching two meters.
Dromaeosaurinae	Indeterminate	A pedal ungual and dentary teeth.	Indeterminate dromaeosaurine remains.
Labocania	L. aguillonae		A teratophonein tyrannosaurine
Ornithomimidae	Indeterminate	Dorsal vertebrae, caudal vertebrae, femora, fibulae, tibiae, metatarsals, ungual manuals, ungual pedals, phalanxes, manual phalanxes, manual phalanges, pedal phalanges, pedal phalanxes, pubes and tarsals.	Indeterminate ornithomimid remains.
Paraxenisaurus	P. normalensis	Caudal vertebrae, femur, peses, manual phalanxes, manual ungual, tarsals, metatarsals, metacarpals, pedal phalanx, pedal phalanges, and pedal unguals.	Represents the first known deinocheirid from North America.
cf. Richardoestesia	Indeterminate	A tooth.	A taxon only known from teeth.
"Saltillomimus"	"S. rapidus"	Partial skeleton.	An informally named ornithomimosaur.
Saurornitholestinae	Indeterminate	A pedal ungual, manual unguals and teeth.	Indeterminate saurornitholestine remains.
Theropoda	Indeterminate	A neural arch, complete and fragmented pedal and manual phalanxes, manual unguals, unspecified vertebra, complete and fragmented caudal vertebrae, chevron, proximal end of femur, and unidentified bone fragments.	Indeterminate theropod remains.
Troodontidae	Indeterminate	A pedal phalanx and a tooth.	Indeterminate troodontid remains.
Tyrannosauridae	Indeterminate	A caudal vertebra, indeterminate vertebrae, tibia, fibula, fragments of ungual phalanges, phalanges, and teeth.	Indeterminate tyrannosaurid remains.

Other Fossils

Pterosaurs

Pterosaurs of the Cerro del Pueblo Formation
Taxa	Presence	Description	Images
Ichnogenus: Pteraichnus	Saltillo, Coahuila	Specimens kept at the Dinosaur Tracks Museum, of the University of Colorado at Denver and the Secretaría de Educación Pública de Coahuila, Mexico

Turtles

Turtles of the Cerro del Pueblo Formation
Genus	Species	Material	Notes	Images
Mexichelys	M. coahuilaensis		New genus for Euclastes coahuilaensis, sea turtle
Chedighaii	C. hutchisoni		A bothremydid turtle
Yelmochelys	Y. rosarioae		A stem-kinosternid turtle

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Other vertebrates

Fish

Fish of the Cerro del Pueblo Formation
Genus	Species	Material	Notes	Images
Lepisosteus	L. sp.
Amiidae	Indet.

Fossil eggs

Mammals

Mammals of the Cerro del Pueblo Formation
Genus	Species	Material	Notes	Images
Cimolomyidae indet.			A cimolomyid multituberculate.
Pediomys	P. cf. elegans		A pediomyid tribosphenidan.
Taeniolabidoidae indet.			A taeniolabidoid multituberculate.
Turgidodon	T. cf. russelli		An alphadontid marsupialiform.

References

^ Weishampel et al., 2004, pp.574-588
^ Fowler, Denver Warwick (2017-11-22). "Revised geochronology, correlation, and dinosaur stratigraphic ranges of the Santonian-Maastrichtian (Late Cretaceous) formations of the Western Interior of North America". PLOS ONE. 12 (11): e0188426. Bibcode:2017PLoSO..1288426F. doi:10.1371/journal.pone.0188426. ISSN 1932-6203. PMC 5699823. PMID 29166406.
^ Ramírez-Velasco, A. A.; Hernández-Rivera, R. (2015). "Diversity of late cretaceous dinosaurs from Mexico" (PDF). Boletín Geológico y Minero. 126 (1): 63–108.
^ Longrich, N.R.; Ramirez Velasco, A.A.; Kirkland, J.; Bermúdez Torres, A.E.; Serrano-Brañas, C.I. (2024). "Coahuilasaurus lipani, a New Kritosaurin Hadrosaurid from the Upper Campanian Cerro Del Pueblo Formation, Northern Mexico". Diversity. 16 (9): 531. doi:10.3390/d16090531.
^ Prieto-Márquez, A. (2013). "Skeletal morphology of Kritosaurus navajovius (Dinosauria:Hadrosauridae) from the Late Cretaceous of the North American south-west, with an evaluation of the phylogenetic systematics and biogeography of Kritosaurini". Journal of Systematic Palaeontology. 12 (2): 133–175. Bibcode:2014JSPal..12..133P. doi:10.1080/14772019.2013.770417. S2CID 84942579.
^ Prieto-Márquez, Albert; Serrano Brañas, Claudia Inés (2012). "Latirhinus uitstlani, a 'broad-nosed' saurolophine hadrosaurid (Dinosauria, Ornithopoda) from the late Campanian (Cretaceous) of northern Mexico". Historical Biology. 24 (6): 607–619. Bibcode:2012HBio...24..607P. doi:10.1080/08912963.2012.671311. S2CID 128964878.
^ Ángel Alejandro Ramírez-Velascoa; Jesús Alvarado-Ortegab (2021). "Review of the taxonomic affinities of Latirhinus uitstlani, an emblematic Mexican hadrosaurid". Journal of South American Earth Sciences. 110. Bibcode:2021JSAES.11003391R. doi:10.1016/j.jsames.2021.103391.
^ Ramírez-Velasco, Á. A.; Aguilar, F. J.; Hernández-Rivera, R.; Gudiño Maussán, J. L.; Rodriguez, M. L.; Alvarado-Ortega, J. (2021). "Tlatolophus galorum, gen. et sp. nov., a parasaurolophini dinosaur from the upper Campanian of the Cerro del Pueblo Formation, Coahuila, northern Mexico". Cretaceous Research. 126: Article 104884. Bibcode:2021CrRes.12604884R. doi:10.1016/j.cretres.2021.104884.
^ Gates, Terry A.; Sampson, Scott D.; Delgado de Jesús, Carlos R.; Zanno, Lindsay E.; Eberth, David; Hernandez-Rivera, René; Aguillón Martínez, Martha C.; Kirkland, James I. (2007). "Velafrons coahuilensis, a new lambeosaurine hadrosaurid (Dinosauria: Ornithopoda) from the Late Campanian Cerro del Pueblo Formation, Coahuila, Mexico". Journal of Vertebrate Paleontology. 27 (4): 917–930. doi:10.1671/0272-4634(2007)27[917:VCANLH]2.0.CO;2. S2CID 86157621.
^ Rivera-Sylva, Héctor E.; Frey, Eberhard; Stinnesbeck, Wolfgang; Amezcua, Natalia; Flores, Flores Huert (2018). "First occurrence of Parksosauridae in Mexico, from the Cerro del Pueblo Formation (Late Cretaceous; late Campanian) at Las Águilas, Coahuila". Boletín de la Sociedad Geológica Mexicana. 70 (3): 779–784. doi:10.18268/BSGM2018v70n3a10. S2CID 202193814.
^ Serrano-Brañas, Claudia Inés; Espinosa-Chávez, Belinda; Maccracken, S. Augusta; Guevara, Daniela Barrera; Torres-Rodríguez, Esperanza (2022-09-30). "First record of caenagnathid dinosaurs (Theropoda, Oviraptorosauria) from the Cerro del Pueblo Formation (Campanian, Upper Cretaceous), Coahuila, Mexico". Journal of South American Earth Sciences. 119: 104046. Bibcode:2022JSAES.11904046S. doi:10.1016/j.jsames.2022.104046. ISSN 0895-9811.
^ Rivera-Sylva, Héctor E.; Longrich, Nicholas R. (2024). "A New Tyrant Dinosaur from the Late Campanian of Mexico Reveals a Tribe of Southern Tyrannosaurs". Fossil Studies. 2 (4): 245–272. doi:10.3390/fossils2040012.
^ Serrano-Brañas, Claudia Inés; Espinosa-Chávez, Belinda; Maccracken, S. Augusta; Gutiérrez-Blando, Cirene; de León-Dávila, Claudio; Ventura, José Flores (April 2020). "Paraxenisaurus normalensis, a large deinocheirid ornithomimosaur from the Cerro del Pueblo Formation (Upper Cretaceous), Coahuila, Mexico". Journal of South American Earth Sciences. 101: 102610. Bibcode:2020JSAES.10102610S. doi:10.1016/j.jsames.2020.102610. S2CID 218968100.
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^ Lockley, M.; Harris, J.D.; and Mitchell, L. 2008. "A global overview of pterosaur ichnology: tracksite distribution in space and time." Zitteliana. B28. p. 187-198. ISSN 1612-4138.
^ James F. Parham; Nicholas D. Pyenson (2010). "New Sea Turtle from the Miocene of Peru and the Iterative Evolution of Feeding Ecomorphologies since the Cretaceous". Journal of Paleontology. 84 (2): 231–247. Bibcode:2010JPal...84..231P. doi:10.1666/09-077R.1. S2CID 62811400.
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^ Don Brinkman; Martha Carolina Aguillon-Martinez; J. Howard Hutchison; Caleb M. Brown (2016). "Yelmochelys rosarioae gen. et sp. nov., a stem kinosternid (Testudines; Kinosternidae) from the Late Cretaceous of Coahuila, Mexico". PaleoBios. 33: 1–20.

Bibliography

Gudiño Maussán, José L.; Aguilar, Felisa J.Hernández; Hernández-Rivera, René (2018). "Aplicación de fotogrametría digital para el registro de las huellas de dinosaurio de la Formación Cerro del Pueblo (Campaniano tardío), Coahuila, México" (PDF). Boletín de la Sociedad Geológica Mexicana. 70 (2): 307–324. doi:10.18268/BSGM2018v70n2a3.