Chitarwata Formation
The Chitarwata Formation is a geological formation in western Pakistan, made up of Oligocene and early Miocene terrestrial fluvial facies. The sediments were deposited in coastal depositional environments (estuarine, strandplain and tidal flats) when Pakistan was partly covered by the Tethys Ocean.
Paleomagnetic data indicates an age range of around 28 to 17 million years ago, with its base in the Oligocene, and its upper boundary, where it meets the overlying Vihowa Formation, of the Early Miocene.
Together with the Vihowa Formation, the Chitarwata Formation records the sedimentation of the Himalayan foreland basin during the collision of the Indian and Asian tectonic plates, the transition from marginal marine to fluvial environments, and the rise of the Himalayas.
Fossil content
The Chitarwata Formation has provided a wealth of terrestrial mammal fossils of the late Paleogene and early Neogene, or Tabenbulakian; the last of the Asian land mammal ages (ALMA).
Among many others, the following fossils are reported from the formation:
Mammals
Artiodactyl
Anthracothere
Anthracotheriidaes of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Anthracotherium | A. adiposum | |||||
A. bugtiense | ||||||
A. silistrense | ||||||
Bugtitherium | B. grandincisivum | |||||
Microbunodon | M. silistrense | |||||
Parabrachyodus | P. hyopotamoides | |||||
Sivameryx | S. palaeindicus |
Entelodonts
Entelodonts of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Paraentelodon | P. sp. |
Giraffid
Giraffids of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Progiraffa | P. exigua |
lophiomerycids
lophiomerycidaes of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Nalameryx | N. savagei | |||||
N. sulaimani | ||||||
Lophibaluchia | L. pilbeami | |||||
Lophiomeryx | L. kargilensis |
Ruminants
Ruminants of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Bugtimeryx | B. pilgrimi | |||||
Paalitherium | P. gurki |
Perissodactyl
Paraceratheriidae
Paraceratheriidaes of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Buzdartherium | B. gulkirao | |||||
Paraceratherium | P. bugtiense |
Rhinocerotidae
Rhinocerotidaes of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Aprotodon | A. smithwoodwardi | |||||
Brachypotherium | B. fatehjangense | |||||
Epiaceratherium | E. magnum | |||||
Pleuroceros | P. blanfordi |
Marsupials
Marsupials of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Asiadidelphis | A. akbarbugtii |
Rodents
Rodents of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Atavocricetodon | A. paaliense | |||||
Baluchimys | B. barryi | |||||
B. chaudryi, | ||||||
B. ganeshapher | ||||||
Bugtimys | B. zafarullahi | |||||
Downsimys | D. margolisi | |||||
Fallomus | F. ginsburgi | |||||
F. quraishyi | ||||||
F. razae | ||||||
Hodsahibia | H. azrae | |||||
H. beamshaiensis | ||||||
H. gracilis | ||||||
Lindsaya | L.derabugtiensis | |||||
Primus | P. cheemai | |||||
P. microps | ||||||
Spanocricetodon | S. sulaiman | |||||
Prokanisamys | P. arifi | |||||
P. kowalskii | ||||||
Pseudocricetodon | P. nawabi | |||||
Welcommoides | W. gurki | |||||
Zindapiria | Z. quadricollis |
Primates
Primates of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Bugtilemur | B. mathesoni | |||||
Bugtipithecus | B. inexpectans | |||||
Guangxilemur | Guangxilemur singsilai | |||||
Phileosimias | P. brahuiorum | |||||
P. kamali |
Proboscidean
Proboscideans of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Deinotherium | D. sp. |
Reptiles
Reptiles of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Asifcroco | A. retrai | A crocodilian. | ||||
Astorgosuchus | A. bugtiensis | A crocodilian. |
Fish
Fishs of the Chitarwata Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Materials | Notes | Image | |
Carcharhinus | C. balochenisis | |||||
C. perseus | ||||||
Cretolamna | Cretolamna twiggensis | |||||
Hemipristis | H. heteropleurus | |||||
Nebrius | N. obliquum |
References
- ^ Chitarwata Formation at Fossilworks.org
Further reading
- S. Adnet, P. -O. Antoine, S. R. Hassan Baqri, J. Crochet, L. Marivaux, J. Welcomme, and G. Métais. 2007. New tropical carcharhinids (chondrichthyes, Carcharhiniformes) from the late Eocene–early Oligocene of Balochistan, Pakistan: Paleoenvironmental and paleogeographic implications. Journal of Asian Earth Sciences 30(2):303-323
- G. Métais, P.-O. Antoine, S. R. H. Baqri, M. Benammi, J.-Y. Crochet, D. Franceschi, L. Marivaux and J.-L. Welcomme. 2006. New remains of the enigmatic cetartiodactyl Bugtitherium grandincisivum Pilgrim, 1908, from the upper Oligocene of the Bugti Hills (Balochistan, Pakistan). Naturwissenschaften 93(7):348-355
- E. H. Lindsay, L. J. Flynn, I. U. Cheema, J. C. Barry, K. Downing, A. R. Rajpar, and S. M. Raza. 2005. Will Downs and the Zinda Pir Dome. Palaeontologia Electronica 8(1):19A:1-18
- L. Marivaux, P.-O. Antoine, S. R. H. Baqri, M. Benammi, and Y. Chaimanee. 2005. Anthropoid primates from the Oligocene of Pakistan (Bugti Hills): Data on early anthropoid evolution and biogeography. Proceedings of the National Academy of Sciences 102(24):8436-8441
- P.-O. Antoine, J.-L. Welcomme, L. Marivaux, I. Baloch, M. Benammi and P. Tassy. 2003. First record of Paleogene Elephantoidea (Mammalia, Proboscidea) from the Bugti Hills of Pakistan. Journal of Vertebrate Paleontology 23(4):977-980
- J.-L. Welcomme and L. Ginsburg. 1997. Mise en évidence de l'Oligocène sur le territoire des Bugti (Balouchistan, Pakistan). Comptes Rendus de l'Académie des Sciences, Série IIA 325(12):999-1004
- L.J. Flynn, L.L. Jacobs, and I.U. Cheema. 1986. Baluchimyinae, A New Ctenodactyloid Rodent Subfamily from the Miocene of Baluchistan. American Museum Novitates 2841:1-58