Kota Formation
Stratigraphy
The lower boundary of the Kota Formation is made of pebbly sandstone, covering the topmost clay seen in the Dharmaram Formation. The Kota Formation has been traditionally divided into 2 main members, the Lower and Upper members, yet more recent work have redivided it into 3. The Lower member can be seen at locations such as Adamilli, Kamavarapukota and Sudikonda, being made of sandstones, with clay clasts, with greater or lower stratification. The Middle Member is well developed along the Continental Gondwana basin, specially towards the northwestern part, and is made of medium to fine white sandstone with clay and concretionary limestone, suggesting the development of paleosols associated with alluvial floodplains. The last member is mostly made of broad sandstone sheets with large clay casts associated with fluvial channels, and has an extension that can be easuly seen on several continuous kilometers. The Uppermost section of the unit is mostly made of limestones and is overlain on an angular unconformity by the Gangapur Formation.
Age
The age of the Kota Formation is controversial. There are no magmatic rocks or volcanic ash beds associated with the Kota Formation, which means that its age cannot be determined directly through radiometric dating. The maximum age of the Kota Formation is constrained by the underlying Upper Dharmaram Formation, which is Early Jurassic, probably Hettangian or Sinemurian, in age. Various researchers have attempted to date the Kota Formation using biostratigraphy. Krishnan (1968), Jain (1973), and Yadagiri and Prasad (1977) favored an Early Jurassic age based on the fish fauna. Govindan (1975) suggested a Middle Jurassic age based on ostracods. In 2006, Bandyopadhyay and Sengupta argued that the fish fauna suggested a Toarcian age for the Upper Kota Formation, possibly extending into the Aalenian, and in turn estimated the Lower Kota to be Sinemurian to Pliensbachian in age. Guntupalli V. R. Prasad, along with various coauthors, has argued for a younger age. In 2001, Vijaya and Prasad proposed based on palynological evidence that the Kota Formation was deposited between the Callovian age of the Middle Jurassic and the Barremian age of the Early Cretaceous. In 2002, Prasad and Manhas argued that the mammal genus Dyskritodon, known only from the Kota Formation and the Early Cretaceous of Morocco, provides evidence for a young age for the Kota Formation. In 2020, Prasad and Parmar argued that the similarity of the dinosaur fauna of the Kota Formation to that of the Middle Jurassic of the United Kingdom supported a Middle Jurassic age for the Kota Formation.
Paleoenvironment
The Kota Formation represents mostly a Continental succession related to a continental rift basin, the Pranhita-Godavari Gondwana Basin of peninsular India. The associated facies of sandstone and limestones are likely related to playa-type lake, with nearby fluvial currents, part of low gradient hanging wall alluvial fans, being deposited on it´s margin. There have been records of freshwater lue green algal stromatolites and oncolites, suggested to be deposited on low energy and low bathymetry lacustrine settings. More recent works have proven the basin hosted in the Early Jurassic a freshwater carbonate wetland marked by the presence of limestones. The environmental model proposed include a depositional cycle marked by several facies types, A for the sublittoral zones of shallow water bodies, followed by palustrine environments, including surfaces with abundance of influence of both plants and animals, specially rhizobrecciation indicating active colonization of the margins by plants, having a similar deposition to the modern Las Tablas de Daimiel wetlands. Associated with the lacustrine facies have recovered microbial bioherms and lacustrine spring mounds, shallow ephemeral ponds with carbonated mud and Phyllopods, pedogenic calcrete under arid seasons and short-lived distributary channels. The depositional setting may have been partially sheltered from the input of siliciclastic materials, except on flooding seasons. Microbial biomats likely developed on shallow waters, while rhizoliths increased it´s presence of abandoned channel fills and pedogenic facies indicate drought seasons.
The carbonate layers with iron-rich grains and mudstones suggest a transition zone between a lake margin and wetland in a rift valley, in a marsh that experienced periodic influxes of iron-rich and barium-rich waters through faults. These waters briefly increased acidity but were neutralized by mixing with alkaline surface water, allowing carbonate formation. Iron-oxidizing microbes likely helped deposit iron oxides, cementing the sediment. During high water flow, these deposits were eroded and transported to deeper waters, forming iron-rich coatings. Thin sandy layers suggest that freshwater influxes occurred, forming carbonate bodies near water discharge points. As the water returned to being more alkaline, carbonates could form again, supporting freshwater Bivalvia, similar to modern geothermal areas of New Zealand.
Fossil content
Color key
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Notes Uncertain or tentative taxa are in small text; |
Ostracoda
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
C. sp. |
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Six carapaces |
A freshwater ostracodan of the Family Pontocyprididae. |
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C. sp. |
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Fourteen incomplete carapaces |
A freshwater ostracodan of the Family Palaeocytheridae. |
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D. cf.sarytirmenensis |
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More than 200 carapaces and valves |
A freshwater ostracodan of the Family Darwinulidae. The most dominant genus locally and the main indicator of both fluvial and lacustrine settings |
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D. kingi |
|
|
Around 120 carapaces and valves |
A freshwater ostracodan of the Family Darwinulidae. |
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D. spp. |
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Nineteen Carapaces |
A freshwater ostracodan of the Family Darwinulidae. |
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E. sp. |
|
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Eight incomplete carapaces |
A freshwater ostracodan of the Family Candoninae. |
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L. spp. |
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Three complete carapaces |
A freshwater ostracodan of the family Limnocytheridae. |
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?S. sp. |
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Single incomplete carapace |
A freshwater ostracodan of the family Cyprididae. |
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T. digitalis |
|
|
Twenty complete carapaces and thirty-six partly broken carapaces. |
A freshwater ostracodan of the family Limnocytheridae. |
Phyllopoda
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
E. alibadadensis |
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Valves |
A freshwater clam shrimp of the family Estheriininae. The most abundant Estheriid in the region and the key element of the Estheriina biozone |
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E. indijurassica |
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Valves |
A freshwater clam shrimp of the family Estheriininae. |
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E. bullata |
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Valves |
A freshwater clam shrimp of the family Estheriininae. |
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E. pranhitaensis |
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Valves |
A freshwater clam shrimp of the family Estheriininae. |
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L. kotaensis |
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Valves |
A freshwater clam shrimp of the family Lioestheriidae. The second key element of the Estheriina biozone |
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L. crustabundis |
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Valves |
A freshwater clam shrimp of the family Lioestheriidae. | ||
L. ssp. |
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Valves |
A freshwater clam shrimp of the family Lioestheriidae. | ||
P. spp. |
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Valves |
A freshwater clam shrimp of the family Estheriininae. |
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P. andhrapradeshia |
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|
Valves |
A freshwater clam shrimp of the family Lioestheriidae. |
Insecta
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
A. shiva |
Tasch outcrop K-2 bed 8 |
|
Right forewing tegmen |
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C. sp. |
Kota limestone ridge |
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Isolated wings |
An Indeterminate Coleopteran. | ||
K. frankmortoni |
Sirpur Taluka, Tasch's K1 outcrop bed 2(A) |
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MCZ 11909, Isolated wing |
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P. lakshmi |
Kota Formation outcrop K-2 |
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MCZ 3046, Isolated wing |
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T. bharataja |
Sirpur Taluka, Tasch's K1 outcrop bed 3(A) |
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No.2013(3013), part and counterpart of well preserved wing |
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T. tulyabhijana |
Sirpur Taluka, Tasch's K1 outcrop bed 2(A) |
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No. 5034, well preserved wing |
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X. alexandri |
Kota Formation outcrop K-2 |
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MCZ 11831, well preserved wing |
Fish
Taxon | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
I. robustus |
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A robust freshwater coelacanth of the family Latimeriidae. Represents the largest member of the local freshwater fauna, measuring up to 70 cm. |
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L. deccanensis |
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A freshwater neopterygian of the family Lepisosteiformes. |
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L. spp. |
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Isolated remains |
A freshwater neopterygian of the family Lepisosteiformes. | ||
L. indicus |
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A freshwater elasmobranch of the family Lonchidiidae. |
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P. egertoni |
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A freshwater neopterygian of the family Dapediidae. |
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P. kingi |
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A freshwater neopterygian of the family Pholidophoridae. |
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P. indicus |
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A freshwater neopterygian of the family Pholidophoridae. |
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P.? sp. |
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A freshwater elasmobranch of the family Polyacrodontidae. |
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Indeterminate |
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A freshwater neopterygian of the family Pycnodontidae, originally classified as Perciformes, yet suggested to be very similar to the Cretaceous pycnodont Stephanodus. |
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Indeterminate |
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|
A freshwater neopterygian of the family Semionotiformes. |
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T. oldhami |
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|
|
A freshwater neopterygian of the family Dapediidae. |
Amphibia
Taxon | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
Indeterminate |
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|
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Indeterminate frog remains, originally referred to Pelobatidae due to be compared with younger Creteaceous Indian frog material |
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Indeterminate |
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Indeterminate caudatan remains, originally referred to Sirenidae due to be compared with younger Creteaceous sirenid material |
Mammaliaforms
Taxon | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
Indeterminate |
Paikasigudem village |
|
Isolated lower molar |
A mammal of the group Australosphenida, resembling the south american genus Asfaltomylos |
||
D.? indicus |
Paikasigudem village |
|
VPL/JU/KM/13, lower left molar |
A dubious mammal of the group Eutriconodonta. This Genus is known from the Early Cretaceous of Morocco, what has been used to suggest a minimum Berrasian age for the Upper Kota Formation |
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G. dattai |
Paikasigudem village |
|
VPL/JU/KM 12 right lower molar |
A mammal of the family Docodontidae |
||
I. pranhitai |
5 km west of Yamanapalli |
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GSI20795, right upper molar |
A mammaliform of the family Morganucodontidae. Includes the informally named "Indozostrodon simpsoni". |
||
I. zofiae |
Paikasigudem village |
|
VPL/JU/KM/20, a left upper premolar |
A mammal described as an eobaatarid multituberculate, but this interpretation has been challenged. |
||
K. haldanei |
5 km west of Yamanapalli |
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GSI19634, right upper molar |
A mammaliform of the family Kuehneotheriidae |
||
N. paikasiensis |
Paikasigudem village |
|
GSI.SR/PAL/12, right upper molar |
A mammal of the family Amphidontidae |
||
P. yadagirii |
Paikasigudem village |
|
VPL/JU/KM/10, right upper molar |
A mammaliaform of the family Morganucodontidae, originally known as "Kotatherium yadagirii" |
||
T. kotaensis |
Paikasigudem village |
|
GSISR/PAL/10, left lower molar |
A mammal of the order Symmetrodonta |
Lepidosauromorpha
Taxon | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
B. rebbanensis |
Paikasigudem village |
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|
A lepidosauromorph originally described as an Iguanian lizard. May actually be a sphenodontian rather than a lizard. |
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G. lateefi |
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|
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A small sphenodontian, with a skull estimated to measure less than 20 mm |
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P. indicus |
Paikasigudem village |
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An indeterminate and dubious lepidosauromorph, originally suggested to be a varanoid lizard |
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R. jaini |
Paikasigudem village |
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A small sphenodont |
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Indeterminate |
Paikasigudem village |
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Distinct from Bharatagama rebbanensis; may include material formerly assigned to the dubious Kota squamate Paikasisaurus indicus. |
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Indeterminate |
Paikasigudem village |
|
|
Indeterminate Sphenodontidae remains |
Testudinata
Taxon | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
I. spatulata |
|
|
|
Mesochelydian stem-turtle, suggested to be related with Condorchelys |
||
Indeterminate |
1 km south of Bodepalli |
|
Carapace fragments |
Indeterminate turtle remains |
Crocodylomorpha
Atoposaurid crocodiles are known from the unit, yet is not clear from what locality.
Taxon | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
Indeterminate |
1 km south of Bodepalli |
Lower Member |
Maxillae, dentaries, teeth |
Indeterminate crocodylomorph remains, previously mixed with thyreophoran material and part of the chimaeric "Andhrasaurus" |
||
Indeterminate |
Kota limestone ridge |
|
Dermal scutes, with a femur and some fragments of other bones |
Indeterminate material referred to crocodylomorphs similar to Teleosaurus |
Pterosauria
Taxon | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
C. indicus |
Kota limestone ridge |
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|
The holotype of Campylognathoides indicus, a pair of premaxillae, may represent a fish rather than a pterosaur. |
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Indeterminate |
|
|
|
Indeterminate pterosaur remains |
Dinosaurs
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
"A. indicus" |
1 km south of Bodepalli |
Lower Member |
Sacral vertebra, vertebral centra, dorsal vertebrae, caudal vertebrae, parts of scapula and ilium, osteoderms |
A chimaera of thyreophoran postcranial material and Crocodylomorph skull pieces. The armor was later suggested to be Ankylosauria indet. And other latter works pointed out it likely belongs to an indeterminate basal thyreophoran. |
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B. tagorei |
|
|
|
A sauropod dinosaur, either a Eusauropoda or more likely a Gravisauria. Represents the best-known Early Jurassic sauropod |
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D. indicus |
Yamanpalli bonebed |
Lower Member |
|
A chimaera of large theropod bones, including and ischium and tooth, probably belonging to a carnosaur, and sauropod bones (two Kotasaurus vertebrae) |
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Indeterminate |
Paikasigudem village |
|
Isolated Teeth |
Five distinct morphotypes have been identified, mostly resemble coelurosaurs or dromaeosauroids. |
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Hypsilophodontidae |
Indeterminate |
Gorlapalli Village |
|
|
Identified originally as a member of Hypsilophodontidae, probably represents a tooth of a basal neornithischian. |
|
K. yamanpalliensis |
Yamanpalli bonebed |
|
Disarticulated remains of at least 12 individuals |
A basal sauropod |
||
Indeterminate |
|
|
|
Indeterminate ornithischian material. Among the teeth, at least five distinct morphotypes have been identified. | ||
R.? spp. |
Paikasigudem village |
|
Isolated Teeth |
Teeth similar to those of the problematic taxon Richardoestesia, of supposed coelurosaur affinities |
||
Indeterminate |
Paikasigudem village |
|
Scute and associated fragmentary limb bones. |
Indeterminate scelidosaurid material. |
||
Sauropoda | Indeterminate. | Yamanpalli bonebed. | • Lower Member | Teeth. | Found in the same bonebed as Kotasaurus, resembles Amygdalodon. | |
Carnosauria | Indeterminate. | Yamanapalli bonebed. | • Lower Member | Teeth. | Found in the same bonebed as Kotasaurus, referred to as |
Megaflora
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
A. kotaense |
Near Kota village |
|
Fossil wood |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
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A. chandrapurensis |
Near Kota village |
|
Fossil wood |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
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A. santacruzense |
Near Kota village |
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Fossil wood |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
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A. pranhitaensis |
Near Kota village |
|
Fossil wood |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
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A. santalense |
Near Kota village |
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Fossil wood |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
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A. spp. |
|
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Fossil wood |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
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A. minutus |
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Branched shoots |
Affinities with Araucariaceae inside Pinales. |
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A. sp. |
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Branched shoots |
Affinities with Araucariaceae inside Pinales. |
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B. expansum |
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Branched shoots |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
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E. conferta |
Near Kota village |
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Branched shoots |
Affinities with Cupressaceae inside Pinales. |
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E. tenerrimus |
Near Kota village |
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Branched shoots |
Affinities with Cupressaceae inside Pinales. |
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E. jabalpurensis |
Near Kota village |
|
Branched shoots |
Affinities with Cupressaceae inside Pinales. |
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E. plana |
Near Kota village |
|
Branched shoots |
Affinities with Cupressaceae inside Pinales. |
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E. sp. |
Near Kota village |
|
Branched shoots |
Affinities with Cupressaceae inside Pinales. |
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C. denticulata |
|
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Isolated fronds |
Affinities with Osmundaceae in the Osmundales. |
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C. indica |
|
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Isolated fronds |
Affinities with Osmundaceae in the Osmundales | ||
C. reversa |
|
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Isolated fronds |
Affinities with Osmundaceae in the Osmundales | ||
C. spp. |
|
|
Isolated fronds |
Affinities with Osmundaceae in the Osmundales | ||
C. kotaense |
Near Kota village |
|
Fossil wood |
Affinities with Podocarpaceae inside Pinales. |
||
C. hymenophylloides |
|
|
Isolated fronds |
Affinities with Polypodiales in the Polypodiidae. Common cosmopolitan Mesozoic fern genus. Recent research has reinterpreted it a stem group of the Polypodiales (closely related to the extant genera Dennstaedtia, Lindsaea, and Odontosoria) |
||
C. sp. |
|
|
Isolated fronds |
Affinities with Polypodiales in the Polypodiidae. | ||
C. kotaense |
Near Kota village |
|
Fossil wood |
Affinities with Cupressaceae inside Pinales. |
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D. falcatus |
|
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Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
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D. kotaense |
|
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Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
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E.rajmahalensis |
|
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Isolated Stems |
Affinities with Equisetaceae inside Equisetales. |
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G. lobata |
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Leafs |
Affinities with Ginkgoaceae inside Ginkgoopsida. |
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G. dixii |
Near Kota village |
|
Fossil wood |
Affinities with Ginkgoaceae inside Ginkgoopsida. |
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H. cf. buchii |
Near Kota village |
|
Isolated pinnae |
Affinities with Dipteridaceae in the Polypodiales. |
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O. vemavarmensis |
|
|
Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
||
P. peregrinum |
Near Kota village |
|
Branched shoots |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
||
P. cf.peregrinum |
Near Kota village |
|
Branched shoots |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
||
P. spp. |
Near Kota village |
|
Branched shoots |
Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. |
||
P. indica |
Near Kota village |
|
Isolated pinnae |
Affinities with Umkomasiaceae in the Pteridospermatophyta. |
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P. mahabalei |
Near Kota village |
|
Fossil wood |
Affinities with Protopinaceae inside Pinales. |
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P. chandrapurensis |
Near Kota village |
|
Fossil wood |
Affinities with Podocarpaceae inside Pinales. |
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P. chiturensis |
Chitur village |
|
Fossil wood |
Affinities with Podocarpaceae inside Pinales. |
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P. krauselii |
Near Kota village |
|
Fossil wood |
Affinities with Podocarpaceae inside Pinales. |
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P. rajmahalense |
Near Kota village |
|
Fossil wood |
Affinities with Podocarpaceae inside Pinales. |
||
P. sewardii |
Near Kota village |
|
Fossil wood |
Affinities with Podocarpaceae inside Pinales. |
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P. sp. |
Near Kota village |
|
Fossil wood |
Affinities with Podocarpaceae inside Pinales. |
||
P. sp. |
|
|
Leaflets |
Broad conifer leaves |
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P. cf. fragilis |
|
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Leaflets |
Affinities with Cycadales in the Cycadopsida. |
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P. fissum |
|
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Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
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P. acutifolium |
|
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Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
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P. cutchense |
|
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Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
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P. cf.sahnii |
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Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
||
P. cf.institacallum |
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Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
||
P. sp. |
|
|
Leaflets |
Affinities with Williamsoniaceae in the Bennettitales. |
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P. sahnii |
Chitur village |
|
Fossil wood |
Affinities with Cupressaceae inside Pinales. |
||
P. liassicum |
Near Kota village |
|
Fossil wood |
Affinities with Cupressaceae inside Pinales. |
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S. kotaensis |
Near Kota village |
|
Isolated Fronds |
Affinities with Dicksoniaceae in the Cyatheales. |
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T. sahnii |
Near Kota village |
|
Fossil wood |
Affinities with Cupressaceae inside Pinales. |
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T. biradarii |
Chitur village |
|
Fossil wood |
Affinities with Cupressaceae inside Pinales. |
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T. antiquum |
Near Kota village |
|
Fossil wood |
Affinities with Cupressaceae inside Pinales. |
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T. sp. |
Near Kota village |
|
Fossil wood |
Affinities with Cupressaceae inside Pinales. |
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T. constricte |
|
|
Branched shoots |
See also
- List of dinosaur-bearing rock formations
- Azilal Formation
- Cañadón Asfalto Formation
- Evergreen Formation
- Mawson Formation
Notes
- ^ It cannot be determined which of the two Kota squamate forms, if either, the holotype of Paikasisaurus indicus pertains to.
References
- ^ Goswami, Suparna; Gierlowski-Kordesch, Elizabeth; Ghosh, Parthasarathi (January 2018). "Sedimentology of the Early Jurassic limestone beds of the Kota Formation: record of carbonate wetlands in a continental rift basin of India". Journal of Paleolimnology. 59 (1): 21–38. Bibcode:2018JPall..59...21G. doi:10.1007/s10933-016-9918-y. ISSN 0921-2728. S2CID 133167210.
- ^ Kutty, T. S.; Jain, S. L.; Chowdhury, T. R. (1987). "Gondwana sequence of the northern Pranhita-Godavari Valley: its stratigraphy and vertebrate faunas" (PDF). Palaeobotanist. 36: 263–282. Retrieved 16 June 2023.
- ^ Lakshminarayana, G. (1994). "Stratigraphy and structural framework of the Gondwana sediments in the Pranhita-Godavari Valley, Andhra Pradesh". Gondwana Nine. 1 (2): 311–330.
- ^ Lakshminarayana, G. (2002). "Evolution in basin fill style during the Mesozoic Gondwana continental break-up in the Godavari Triple junction, SE India". Gondwana Research. 5 (1): 227–244. Bibcode:2002GondR...5..227L. doi:10.1016/S1342-937X(05)70906-0. Retrieved 16 June 2023.
- ^ Sengupta, S. (2003). "Gondwana sedimentation in the Pranhita–Godavari Valley: a review". Journal of Asian Earth Sciences. 21 (6): 633–642. Bibcode:2003JAESc..21..633S. doi:10.1016/S1367-9120(02)00052-4. Retrieved 16 June 2023.
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