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  • 21 Aug, 2019

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List Of Scandentians

Scandentia is an order of small mammals. Members of this order are called scandentians, or treeshrews. They are primarily found in Southeast Asia, with the Madras treeshrew instead in mainland India. They range in size from the Bornean smooth-tailed treeshrew, at 11 cm (4 in) plus a 9 cm (4 in) tail, to the striped treeshrew, at 23 cm (9 in) plus a 13 cm (5 in) tail. They generally live in forests, and primarily eat insects and fruit. No population estimates have been made for any scandentian species, though the Nicobar treeshrew is categorized as endangered.

The twenty-three extant species of Scandentia are grouped into two families: Tupaiidae, which contains twenty-two species within three genera, and Ptilocercidae, which contains a single species, the pen-tailed treeshrew. Only a few extinct scandentian species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.

Conventions

IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically Endangered (0 species)
 EN Endangered (1 species)
 VU Vulnerable (1 species)
 NT Near threatened (0 species)
 LC Least concern (15 species)
Other categories
 DD Data deficient (6 species)
 NE Not evaluated (0 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the scandentian's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "†".

Classification

The order Scandentia consists of two families: Tupaiidae, which contains twenty-two species within three genera, and Ptilocercidae, which contains a single species. Many of these species are further subdivided into subspecies. This does not include hybrid species or extinct prehistoric species.

Family Ptilocercidae

Family Tupaiidae

  • Genus Anathana (Madras treeshrew): one species
  • Genus Dendrogale (smooth-tailed treeshrews): two species
  • Genus Tupaia (treeshrews): nineteen species
Scandentia  

Scandentians

The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis.

Family Ptilocercidae

Genus Ptilocercus Gray, 1848 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Pen-tailed treeshrew

Drawing of brown treeshrew

P. lowii
Gray, 1848

Two subspecies
  • P. l. continentis
  • P. l. lowii
Southeast Asia
Map of range
Size: 13–14 cm (5–6 in) long, plus 16–19 cm (6–7 in) tail

Habitat: Forest

Diet: Insects and fruit
 LC 


Unknown Population declining

Family Tupaiidae

Genus Anathana Lyon, 1913 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Madras treeshrew

Brown treeshrew

A. ellioti
(Waterhouse, 1850)
India
Map of range
Size: 16–19 cm (6–7 in) long, plus 16–20 cm (6–8 in) tail

Habitat: Forest and shrubland

Diet: Insects, other invertebrates, and fruit
 LC 


Unknown Population declining

Genus Dendrogale Gray, 1848 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bornean smooth-tailed treeshrew


D. melanura
(Thomas, 1892)

Two subspecies
  • D. m. baluensis
  • D. m. melanura
Northern Borneo
Map of range
Size: 11–15 cm (4–6 in) long, plus 9–14 cm (4–6 in) tail

Habitat: Forest

Diet: Fruit and insects
 DD 


Unknown Population declining

Northern smooth-tailed treeshrew

Brown treeshrew

D. murina
(Schlegel, Müller, 1843)
Mainland Southeast Asia
Map of range
Size: About 12 cm (5 in) long, plus 4–5 cm (2 in) tail

Habitat: Forest

Diet: Fruit, arthropods, and small vertebrates
 LC 


Unknown Population declining

Genus Tupaia Raffles, 1821 – nineteen species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bangka Island treeshrew


T. discolor
Lyon, 1906
Bangka Island, Indonesia Size: 13–21 cm (5–8 in) long, plus 12–20 cm (5–8 in) tail

Habitat: Unknown

Diet: Fruit and insects
 DD 


Unknown Unknown

Common treeshrew

Gray/brown treeshrew

T. glis
(Diard, 1820)
Southeast Asia
Map of range
Size: 13–21 cm (5–8 in) long, plus 12–20 cm (5–8 in) tail

Habitat: Forest

Diet: Insects, other arthropods, and fruit
 LC 


Unknown Population declining

Golden-bellied treeshrew


T. chrysogaster
Miller, 1903
Mentawai Islands, Indonesia
Map of range
Size: 12–21 cm (5–8 in) long, plus 14–20 cm (6–8 in) tail

Habitat: Forest

Diet: Fruit and insects
 VU 


Unknown Unknown

Horsfield's treeshrew

Brown treeshrew

T. javanica
Horsfield, 1822
Western Indonesia
Map of range
Size: 12–21 cm (5–8 in) long, plus 14–20 cm (6–8 in) tail

Habitat: Forest

Diet: Fruit and insects
 LC 


Unknown Population declining

Javan treeshrew


T. hypochrysa
Thomas, 1895
Java, Indonesia Size: 13–21 cm (5–8 in) long, plus 12–20 cm (5–8 in) tail

Habitat: Unknown

Diet: Fruit and insects
 DD 


Unknown Unknown

Kalimantan treeshrew


T. salatana
Lyon, 1913
Southern Borneo, Indonesia Size: 12–21 cm (5–8 in) long, plus 14–20 cm (6–8 in) tail

Habitat: Forest

Diet: Termites and ants
 DD 


Unknown Unknown

Large treeshrew

Stuffed brown treeshrew

T. tana
Raffles, 1821

Fifteen subspecies
  • T. t. banguei
  • T. t. besara
  • T. t. bunoae
  • T. t. cervicalis
  • T. t. chrysura
  • T. t. kelabit
  • T. t. kretami
  • T. t. lingae
  • T. t. masae
  • T. t. nitida
  • T. t. paitana
  • T. t. sirhassenensis
  • T. t. speciosa
  • T. t. tana
  • T. t. utara
Southeast Asia
Map of range
Size: About 22 cm (9 in) long, plus 17 cm (7 in) tail

Habitat: Forest

Diet: Insects and other invertebrates, as well as fruit
 LC 


Unknown Population declining

Long-footed treeshrew

Stuffed brown treeshrew

T. longipes
Thomas, 1893
Borneo
Map of range
Size: 12–21 cm (5–8 in) long, plus 14–20 cm (6–8 in) tail

Habitat: Forest

Diet: Fruit and insects
 LC 


Unknown Population declining

Mindanao treeshrew


T. everetti
Thomas, 1892
Mindanao, Philippines
Map of range
Size: 17–22 cm (7–9 in) long, plus 11–18 cm (4–7 in) tail

Habitat: Forest

Diet: Small animals, insects, fruit, vegetables, and eggs
 LC 


Unknown Population steady

Mountain treeshrew


T. montana
Thomas, 1892

Two subspecies
  • T. m. baluensis
  • T. m. montana
Northern Borneo
Map of range
Size: About 20 cm (8 in) long, plus 18 cm (7 in) tail

Habitat: Forest

Diet: Omnivorous
 LC 


Unknown Unknown

Nicobar treeshrew

Gray and brown treeshrew

T. nicobarica
(Zelebor, 1869)

Two subspecies
  • T. n. nicobarica
  • T. n. surda
Nicobar Islands, India
Map of range
Size: About 14 cm (6 in) long, plus 18 cm (7 in) tail

Habitat: Forest

Diet: Insects as well as fruit
 EN 


Unknown Population declining

Northern treeshrew

Gray and brown treeshrew

T. belangeri
(Wagner, 1841)

Two subspecies
  • T. b. belangeri
  • T. b. chinensis
Southeast Asia
Map of range
Size: 12–21 cm (5–8 in) long, plus 14–20 cm (6–8 in) tail

Habitat: Forest and shrubland

Diet: Insects as well as fruit
 LC 


Unknown Population steady

Painted treeshrew

Red and brown treeshrew

T. picta
Thomas, 1892

Two subspecies
  • T. p. fuscior
  • T. p. picta
Borneo
Map of range
Size: About 21 cm (8 in) long, plus 15 cm (6 in) tail

Habitat: Forest

Diet: Fruit and insects
 LC 


Unknown Population declining

Palawan treeshrew


T. palawanensis
Thomas, 1894
Palawan, Philippines
Map of range
Size: 12–21 cm (5–8 in) long, plus 14–20 cm (6–8 in) tail

Habitat: Forest

Diet: Insects and fruit
 LC 


Unknown Population steady

Pygmy treeshrew

Two stuffed brown treeshrews

T. minor
Günther, 1876

Four subspecies
  • T. m. humeralis
  • T. m. malaccana
  • T. m. minor
  • T. m. sincipis
Southeast Asia
Map of range
Size: 11–14 cm (4–6 in) long, plus 13–17 cm (5–7 in) tail

Habitat: Forest

Diet: Small animals, fruit leaves, seeds, and carrion
 LC 


Unknown Population declining

Ruddy treeshrew

Drawing of brown treeshrew

T. splendidula
Gray, 1865

Five subspecies
  • T. s. carimatae
  • T. s. lucida
  • T. s. natunae
  • T. s. riabus
  • T. s. splendidula
Southern Borneo
Map of range
Size: About 18 cm (7 in) long, plus 14 cm (6 in) tail

Habitat: Forest

Diet: Fruit and insects
 LC 


Unknown Population declining

Slender treeshrew


T. gracilis
Thomas, 1893

Three subspecies
  • T. g. edarata
  • T. g. gracilis
  • T. g. inflata
Borneo and nearby islands
Map of range
Size: 12–21 cm (5–8 in) long, plus 14–20 cm (6–8 in) tail

Habitat: Forest

Diet: Fruit and insects
 LC 


Unknown Population declining

Striped treeshrew

Drawing of brown treeshrew

T. dorsalis
Schlegel, 1857
Borneo
Map of range
Size: 14–23 cm (6–9 in) long, plus 10–13 cm (4–5 in) tail

Habitat: Forest

Diet: Insects, fruit, seeds, and leaves
 DD 


Unknown Population declining

Sumatran treeshrew


T. ferruginea
Raffles, 1821
Sumatra and Tanahbala, Indonesia Size: 13–21 cm (5–8 in) long, plus 12–20 cm (5–8 in) tail

Habitat: Unknown

Diet: Fruit and insects
 DD 


Unknown Unknown

References

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Sources