The
Quseir Formation is a
geological formation in the vicinity of the
Kharga Oasis in
Egypt . It is
Campanian in age. The lithology largely consists of soft shale with hard bands of
sandstone ,
siltstone and
phosphorite . The environment of deposition was nearshore to freshwater fluvio-lacustrine characterized by moist and aquatic habitats with a tropical warm-humid climate. It is conformably overlain by the marine late Campanian-
Maastrichtian Duwi Formation, and unconformably overlies the
Turonian Taref Formation. The sauropod dinosaurs
Mansourasaurus and
Igai are known from the formation, as well as the proximal fibula of an indeterminate theropod and possible remains tentively assigned to
Spinosaurus . Additionally the lungfish genera
Lavocatodus and Protopterus , the crocodyliform
Wahasuchus and the
bothremydid turtle
Khargachelys are also known.
Fossil content
Color key
Notes Uncertain or tentative taxa are in small text ; crossed out taxa are discredited.
Dinosaurs
Theropods including a proximal fibula and specimens tentatively assigned to Cenomanian theropod genera Spinosaurus and Carcharodontosaurus are now classified as Theropoda indet.
Sauropods
Sauropods of the Quseir Formation
Genus
Species
Location
Material
Notes
Images
Mansourasaurus
M. shahinae
a fragment of the skull roof , a part of the lower braincase , the dentaries of the lower jaws, three neck vertebrae , two back vertebrae , eight ribs , the right scapula , the right coracoid , both humeri , a radius , a third metacarpal , three metatarsals , and parts of osteoderms .
A lithostrotian sauropod
Igai
I. semkhu
A saltosaurid titanosaur
Fish
Boney Fish
Fish of the Quseir Formation
Genus
Species
Location
Material
Notes
Images
Lavocatodus
L. giganteus
L.? humei
L.? protopteroides
Protopterus
P. nigeriensis
Protopterus_dolloi_Boulenger2 A modern Protopterus species
Cartilaginous Fish
Crocodyliformes of the Quseir Formation
Genus
Species
Location
Material
Notes
Images
Wahasuchus
W. egyptensis
Fossils of skull and jaw fragments, dorsal vertebrae, and fragmentary appendicular remains
A mesoeucrocodylian
Turtles
References
^ Quseir Formation at Fossilworks .org
^ Mahmoud, Magdy S. (April 2003). "Palynology and palaeoenvironment of the Quseir Formation (Campanian) from central Egypt". Journal of African Earth Sciences . 36 (3): 135– 148. Bibcode :2003JAfES..36..135M . doi :10.1016/s0899-5362(03)00047-2 . ISSN 1464-343X .
^ Sallam, Hesham M.; O'Connor, Patrick M.; Kora, Mahmoud; Sertich, Joseph J.W.; Seiffert, Erik R.; Faris, Mahmoud; Ouda, Khaled; El-Dawoudi, Iman; Saber, Sara (May 2016). "Vertebrate paleontological exploration of the Upper Cretaceous succession in the Dakhla and Kharga Oases, Western Desert, Egypt" . Journal of African Earth Sciences . 117 : 223– 234. Bibcode :2016JAfES.117..223S . doi :10.1016/j.jafrearsci.2016.01.022 . ISSN 1464-343X .
^ Sallam, Hesham M.; Gorscak, Eric; O’Connor, Patrick M.; El-Dawoudi, Iman A.; El-Sayed, Sanaa; Saber, Sara; Kora, Mahmoud A.; Sertich, Joseph J. W.; Seiffert, Erik R. (2018-01-29). "New Egyptian sauropod reveals Late Cretaceous dinosaur dispersal between Europe and Africa". Nature Ecology & Evolution . 2 (3): 445– 451. Bibcode :2018NatEE...2..445S . doi :10.1038/s41559-017-0455-5 . ISSN 2397-334X . PMID 29379183 . S2CID 3375335 .
^ Gorscak, Eric; Lamanna, Matthew C.; Schwarz, Daniela; Díaz, Verónica díez; Salem, Belal S.; Sallam, Hesham M.; Wiechmann, Marc Filip (2023-07-20). "A new titanosaurian (Dinosauria: Sauropoda) from the Upper Cretaceous (Campanian) Quseir Formation of the Kharga Oasis, Egypt" . Journal of Vertebrate Paleontology . 42 (6). doi :10.1080/02724634.2023.2199810 . ISSN 0272-4634 .
^ Salem, Belal S.; O'Connor, Patrick M.; Gorscak, Eric; El-Sayed, Sanaa; Sertich, Joseph J.W.; Seiffert, Erik; Sallam, Hesham M. (2021-07-01). "Dinosaur remains from the Upper Cretaceous (Campanian) of the Western Desert, Egypt" . Cretaceous Research . 123 : 104783. Bibcode :2021CrRes.12304783S . doi :10.1016/j.cretres.2021.104783 . ISSN 0195-6671 . S2CID 233900405 .
^ Churcher, C. S.; De iuliis, G. (2001). "A new species of Protopterus and a revision of Ceratodus humei (Dipnoi: Ceratodontiformes) from the Late Cretaceous Mut Formation of eastern Dakhleh Oasis, Western Desert of Egypt" . Palaeontology . 44 (2): 305– 323. Bibcode :2001Palgy..44..305C . doi :10.1111/1475-4983.00181 . ISSN 0031-0239 .
^ Augustin, Felix J.; Hartung, Josephina; Kampouridis, Panagiotis (2023), Hamimi, Zakaria; Khozyem, Hassan; Adatte, Thierry; Nader, Fadi H. (eds.), "Dinosaur Faunas of Egypt—The Terrestrial Late Cretaceous Vertebrate Record" , The Phanerozoic Geology and Natural Resources of Egypt , Cham: Springer International Publishing, pp. 253– 284, doi :10.1007/978-3-030-95637-0_9 , ISBN 978-3-030-95636-3 , retrieved 2024-11-26
^ Claeson, Kerin M; Sallam, Hesham M; O'Connor, Patrick M.; Sertich, Joseph J. W. (2014). "A revision of the Upper Cretaceous lepidosirenid lungfishes from the Quseir Formation, Western Desert, central Egypt". Journal of Vertebrate Paleontology . 34 (4): 760– 766. Bibcode :2014JVPal..34..760C . doi :10.1080/02724634.2014.838574 . S2CID 17120150 .
^ Saber, Sara; Sertich, Joseph J.W.; Sallam, Hesham M.; Ouda, Khaled A.; O'Connor, Patrick M.; Seiffert, Erik R. (October 2018). "An enigmatic crocodyliform from the Upper Cretaceous Quseir Formation, central Egypt" . Cretaceous Research . 90 : 174– 184. Bibcode :2018CrRes..90..174S . doi :10.1016/j.cretres.2018.04.004 . ISSN 0195-6671 .
^ AbdelGawad, Mohamed; Pérez-García, Adán; Hirayama, Ren; Mohesn, Sara; Tantawy, Abdel-Aziz; Abu El-Kheir, Gebely (February 2023). "The First Side-Necked Turtle (Pleurodira, Bothremydidae) from the Campanian (Late Cretaceous) of Egypt" . Diversity . 15 (2): 284. doi :10.3390/d15020284 . ISSN 1424-2818 .
^ Augustin, Felix J.; Hartung, Josephina; Kampouridis, Panagiotis (2023), Hamimi, Zakaria; Khozyem, Hassan; Adatte, Thierry; Nader, Fadi H. (eds.), "Dinosaur Faunas of Egypt—The Terrestrial Late Cretaceous Vertebrate Record" , The Phanerozoic Geology and Natural Resources of Egypt , Cham: Springer International Publishing, pp. 253– 284, doi :10.1007/978-3-030-95637-0_9 , ISBN 978-3-030-95636-3 , retrieved 2024-10-18
^ Sallam, Hesham M.; Gorscak, Eric; O’Connor, Patrick M.; El-Dawoudi, Iman A.; El-Sayed, Sanaa; Saber, Sara; Kora, Mahmoud A.; Sertich, Joseph J. W.; Seiffert, Erik R.; Lamanna, Matthew C. (2018-01-29). "New Egyptian sauropod reveals Late Cretaceous dinosaur dispersal between Europe and Africa" . Nature Ecology & Evolution . 2 (3): 445– 451. Bibcode :2018NatEE...2..445S . doi :10.1038/s41559-017-0455-5 . ISSN 2397-334X . PMID 29379183 .
^ Gorscak, Eric; Lamanna, Matthew C.; Schwarz, Daniela; Díaz, Verónica Díez; Salem, Belal S.; Sallam, Hesham M.; Wiechmann, Marc Filip (2022-12-14). "A new titanosaurian (Dinosauria: Sauropoda) from the Upper Cretaceous (Campanian) Quseir Formation of the Kharga Oasis, Egypt" . Journal of Vertebrate Paleontology . 42 (6). doi :10.1080/02724634.2023.2199810 . ISSN 0272-4634 .
^ Saber, Sara; Sertich, Joseph J.W.; Sallam, Hesham M.; Ouda, Khaled A.; O'Connor, Patrick M.; Seiffert, Erik R. (2018). "An enigmatic crocodyliform from the Upper Cretaceous Quseir Formation, central Egypt" . Cretaceous Research . 90 : 174– 184. Bibcode :2018CrRes..90..174S . doi :10.1016/j.cretres.2018.04.004 .